ECOS species

Tree Species

In 1988, the 12 treatments consisted of 11 tree species, and one 'control', in which regeneration would occur naturally. Trees in the surrounding plots soon overtopped the 'control', so it did not prove valuable. Two exotic species have been removed to prevent their spread, and three species have either died or did not establish well.

As of 2003, six of the 11 originally planted species have survived and now form closed canopies. These six species differ in traits such as N-fixation, rooting depth, and growth rates. The following table is based on Fisher (1995), Haggar et al. (1997), and our measurements in 2003:

Species	Family	Traits	Mean tree size (age 15 y)
			dbh (cm)	height (m)
Hyeronima alchorneoides	Euphorbiaceae	heavy litter	22.0	21.4
Pentaclethra macroloba	Fabaceae	N fixer	15.8	8.5
Pinus patula (exotic)	Pinaceae	deep roots	28.6	22.0
Virola koschnyi	Myristicaceae	deep roots	21.4	18.0
Vochysia ferruginea	Vochysiaceae		34.2	25.5
Vochysia guatemalensis	Vochysiaceae	deep roots	28.4	22.3

REFERENCES

The references for the tree species pdf attachments are:

Dvorak, W.S. 2002. Pinus tecunumanii Eguiluz & J.P. Perry. IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 514-517.

Flores, E.M. 2002. Hyeronima alchorneoides Allemao IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 514-517.

Flores, E.M. 2002. Pentaclethra macroloba (Willd.) Kuntze. IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 601-604.

Flores, E.M. 2002. Virola koschnyi Warb. IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 772-774.

Fournier, L.A. 2002. Vochysia guatemalensis Donn. Sm. IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 778-780.

Rodríguez Sánchez, L. and E. Müller. 2002. Vochysia ferruginea Mart. IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 775-777.

References

Flores, E.M. 2002 IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 514-517.

Flores, E.M. 2002. Chapter I. Seed biology. IN: Vozzo, J. A. (ed.). Tropical Tree Seed Manual. Agricultural Handbook 721. Washington: Forest Service, United States Department of Agriculture. p. 13-118.

Flores, E. M. 2004 Tree Seed Physiology. IN: Burley, J., Evans, J. & Youngquist, J. (eds.). Encyclopedia of Forest Sciences. Oxford, UK: Elsevier Ltd. P 1590-1600.

Flores, E.M; Obando, G. 2003 Arboles del trópico Húmedo. Valorsocioeconómico. Cartago, Costa Rica: Editorial Tecnológica de Costa Rica. 922 p. (There is a related data base in www.fundecor.org ).